{"id":35169,"date":"2020-04-15T00:11:49","date_gmt":"2020-04-14T16:11:49","guid":{"rendered":"https:\/\/case.ntu.edu.tw\/blog\/?p=35169"},"modified":"2023-06-06T11:05:25","modified_gmt":"2023-06-06T03:05:25","slug":"%e6%b5%b7%e6%b4%8b%e5%be%ae%e7%94%9f%e7%89%a9%e7%9f%a5%e5%a4%9a%e5%b0%91-%e8%87%ba%e7%81%a3%e6%b5%b7%e6%b4%8b%e7%a7%91%e5%ad%b8%e7%a0%94%e7%a9%b6%ef%bc%9a%e5%8d%8a%e4%b8%96%e7%b4%80%e7%9a%84","status":"publish","type":"post","link":"https:\/\/case.ntu.edu.tw\/blog\/?p=35169","title":{"rendered":"\u3010\u79d1\u5b78\u53f2\u6c99\u9f8d\u3011\u3008\u6d77\u6d0b\u5fae\u751f\u7269\u77e5\u591a\u5c11\u3009&#038;\u3008\u81fa\u7063\u6d77\u6d0b\u79d1\u5b78\u7814\u7a76\uff1a\u534a\u4e16\u7d00\u7684\u98a8\u83ef\u3009"},"content":{"rendered":"<p style=\"text-align: center;\"><iframe src=\"https:\/\/www.youtube.com\/embed\/cOIkSt1cbFs\" width=\"560\" height=\"315\" frameborder=\"0\" allowfullscreen=\"allowfullscreen\"><\/iframe><\/p>\n<p><span style=\"color: #800000;\"><strong>\u3008\u6d77\u6d0b\u5fae\u751f\u7269\u77e5\u591a\u5c11\u3009<\/strong><\/span><\/p>\n<p><span style=\"color: #800080;\"><strong>\u6d77\u6d0b\u5fae\u751f\u7269\u7a2e\u985e\u7e41\u591a\uff0c\u5305\u62ec\u81ea\u71df\u7d30\u83cc\u3001\u7570\u71df\u7d30\u83cc\u3001\u53e4\u7d30\u83cc\u3001\u75c5\u6bd2\u3001\u7e96\u6bdb\u87f2\u3001\u97ad\u6bdb\u87f2\u3001\u771f\u83cc\u3001\u85fb\u985e\u7b49\u7b49\uff0c\u6700\u65e9\u53ef\u8ffd\u6eaf\u5230 35 \u5104\u5e74\u524d\u7684\u7d30\u83cc\u5316\u77f3\u3002\u672c\u8b1b\u6b21\u56de\u9867\u6d77\u6d0b\u5fae\u751f\u7269\u63a2\u7d22\u53f2\u7684\u91cd\u8981\u91cc\u7a0b\u7891\uff0c\u4ee5\u53ca\u76ee\u524d\u5df2\u78ba\u5b9a\u7684\u4e00\u4e9b\u95dc\u65bc\u6d77\u6d0b\u5fae\u751f\u7269\u7684\u4e8b\u5be6\u3002<\/strong><\/span><\/p>\n<p><strong>\u8b1b\u5e2b\uff1a\u8b1d\u5fd7\u8c6a\uff5c\u81fa\u7063\u5927\u5b78\u6d77\u6d0b\u7814\u7a76\u6240\u6559\u6388<\/strong><\/p>\n<p>\u7d30\u83cc\u662f\u6d77\u6d0b\u4e2d\u91cd\u8981\u7684\u5206\u89e3\u8005\uff0c\u4f46\u662f\u9664\u6b64\u4e4b\u5916\uff0c\u4eba\u5011\u5c0d\u65bc\u6d77\u6d0b\u7d30\u83cc\u7684\u7d55\u5927\u90e8\u5206\u4e86\u89e3\uff0c\u90fd\u662f\u8fd1 50 \u5e74\u624d\u9678\u7e8c\u5efa\u7acb\u8d77\u4f86\u7684\u3002\u9996\u5148\u662f\u6d77\u6d0b\u7d30\u83cc\u5438\u6536\u6709\u6a5f\u7269\u7684\u6548\u7387\uff0c\u662f\u5728 1970 \u5e74\u4ee3\u5229\u7528\u653e\u5c04\u6027\u540c\u4f4d\u7d20 C14 \uff0c\u6a19\u5b9a\u5404\u7a2e\u6709\u6a5f\u7269\u57f9\u990a\u5be6\u9a57\uff0c\u624d\u78ba\u5b9a\u7d30\u83cc\u5927\u7d04\u53ef\u5438\u6536 70% \u7684\u6709\u6a5f\u7269\uff1b\u6d77\u6d0b\u4e2d\u7684\u7d30\u83cc\u6578\u91cf\u540c\u6a23\u662f\u5728 1970 \u5e74\u4ee3\uff0c\u85c9\u7531\u87a2\u5149\u986f\u5fae\u93e1\u78ba\u8a8d\u4e00\u516c\u5347\u7684\u6d77\u6c34\uff0c\u5927\u7d04\u542b\u6709 10 \u5104\u500b\u7d30\u83cc\u3002\u7d30\u83cc\u7684\u751f\u9577\u901f\u7387\u5247\u662f\u5230\u4e86 1980 \u5e74\u4ee3\uff0c\u5229\u7528\u653e\u5c04\u6027\u540c\u4f4d\u7d20\u4e09\u539f\u5b50\u6c2b\uff0c\u6a19\u5b9a\u7d30\u83cc\u751f\u9577\u6240\u9700\u7684 DNA \u539f\u6599\uff1b\u53e6\u5916\u5229\u7528\u57f9\u990a\u5be6\u9a57\uff0c\u6e2c\u91cf\u7d30\u83cc\u6d88\u8017\u7684\u6c27\u6c23\u91cf\uff0c\u4f30\u7b97\u7d30\u83cc\u7684\u547c\u5438\u7387\uff0c\u63a8\u7b97\u51fa\u7d30\u83cc\u7684\u4ee3\u8b1d\u7387\u4f54\u7cfb\u7d71\u6574\u9ad4\u4ee3\u8b1d\u7387\u8d85\u904e 50% \u3002<\/p>\n<p>\u5728\u7f3a\u4e4f\u71df\u990a\u7684\u74b0\u5883\u4e2d\u4e5f\u80fd\u751f\u9577\u5f97\u5f88\u597d\u7684\u85cd\u7da0\u83cc\uff0c\u662f\u6d77\u6d0b\u4e3b\u8981\u7684\u57fa\u790e\u751f\u7522\u8005\uff0c\u4f54\u6d77\u6d0b\u6574\u9ad4\u751f\u7522\u91cf\u7684 60% \u3002\u97ad\u6bdb\u87f2\u5247\u662f\u7d30\u83cc\u7684\u4e3b\u8981\u6355\u98df\u8005\uff0c\u5b83\u540c\u6642\u4e5f\u6703\u88ab\u6d6e\u6e38\u52d5\u7269\u6355\u98df\uff0c\u56e0\u6b64\u5f62\u6210\u4e00\u500b\u6d77\u6d0b\u5fae\u751f\u7269\u98df\u7269\u5708 (microbial loop) \u3002\u5230\u4e86 1980 \u5e74\u4ee3\u672b\u671f\uff0c\u79d1\u5b78\u5bb6\u5229\u7528\u6d41\u5f0f\u7d30\u80de\u5100\u8ddf\u96fb\u5b50\u986f\u5fae\u93e1\uff0c\u4e0d\u4f46\u767c\u73fe\u4e86\u4e16\u754c\u4e0a\u6700\u5c0f\u7684\u85cd\u7da0\u83cc\u539f\u7da0\u7403\u85fb\uff0c\u9084\u767c\u73fe\u4e00\u516c\u5347\u7684\u6d77\u6c34\u5927\u7d04\u542b\u6709 100 \u5104\u500b\u75c5\u6bd2\uff0c\u9019\u4e9b\u5e73\u6642\u8655\u65bc\u4f11\u7720\u72c0\u614b\u7684\u75c5\u6bd2\uff0c\u4e0d\u6642\u6703\u6d3b\u5316\u8d77\u4f86\uff0c\u628a\u7d30\u83cc\u88c2\u89e3\u6210\u6709\u6a5f\u7269\uff0c\u5f9e\u800c\u884d\u751f\u51fa\u75c5\u6bd2\u5708 (viral loop) \u7684\u6982\u5ff5\u3002\u6d77\u6d0b\u5fae\u751f\u7269\u98df\u7269\u7db2\u7684\u60f3\u6cd5\u8207\u7d50\u69cb\uff0c\u9019\u624d\u5b8c\u6574\u5730\u5efa\u7acb\u8d77\u4f86\u3002<\/p>\n<p>\u5927\u90e8\u5206\u7684\u6d77\u6d0b\u5fae\u751f\u7269\u90fd\u7121\u6cd5\u57f9\u990a\uff0c\u60f3\u8981\u77e5\u9053\u6d77\u6d0b\u88e1\u6709\u591a\u5c11\u5fae\u751f\u7269\uff0c\u5404\u6709\u4ec0\u9ebc\u529f\u80fd\uff0c\u53c8\u8981\u5982\u4f55\u8457\u624b\u505a\u7814\u7a76\uff1f\u9019\u5c31\u662f\u5206\u5b50\u751f\u7269\u5b78\u767c\u63ee\u64c5\u5834\u4e4b\u8655\u4e86\uff0c\u8209\u51e1 16S RNA \u57fa\u56e0\u5e8f\u5217\u5206\u6790\u3001\u57fa\u56e0\u8907\u88fd\u5eab\u7814\u7a76\u3001\u6a19\u7684\u57fa\u56e0\u9ad8\u901a\u91cf\u5b9a\u5e8f\uff0c\u5206\u5b50\u751f\u7269\u5b78\u5bb6\u796d\u51fa\u5404\u7a2e\u7814\u7a76\u624b\u6cd5\uff0c\u9010\u6f38\u5f97\u51fa\u6d77\u6d0b\u7d30\u83cc\u7684\u591a\u6a23\u6027\u5f88\u9ad8\uff0c\u4f46\u662f\u5927\u90e8\u5206\u83cc\u7a2e\u7684\u6578\u91cf\u90fd\u5f88\u5c11\u7684\u4e8b\u5be6\u3002\u6d77\u6d0b\u7d30\u83cc\u5c0d\u65bc\u751f\u7269\u5730\u7403\u5316\u5b78\u7684\u529f\u80fd\uff0c\u5247\u662f\u53cd\u6620\u5728\u6d77\u6d0b\u7684\u78b3\u3001\u6c2e\u3001\u78f7\u7b49\u7b49\u5143\u7d20\u7684\u5faa\u74b0\uff0c\u5c0d\u65bc\u6d77\u6d0b\u9178\u5316\u4ee5\u53ca\u5730\u7403\u6574\u9ad4\u6c23\u5019\uff0c\u5177\u6709\u95dc\u9375\u6027\u7684\u5f71\u97ff\u3002\u9019\u662f\u5c6c\u65bc\u7bc4\u570d\u66f4\u5ee3\u7684\u751f\u614b\u7cfb\u7d71\u6d77\u6d0b\u5b78\u7814\u7a76\u9818\u57df\uff0c\u4e0d\u904e\u90a3\u5c31\u662f\u53e6\u4e00\u500b\u6709\u5f85\u4f86\u65e5\u7684\u6545\u4e8b\u4e86\u3002<\/p>\n<p>&nbsp;<\/p>\n<p><span style=\"color: #800000;\"><strong>\u3008\u81fa\u7063\u6d77\u6d0b\u79d1\u5b78\u7814\u7a76\uff1a\u534a\u4e16\u7d00\u7684\u98a8\u83ef\u3009<\/strong><\/span><\/p>\n<p><span style=\"color: #800080;\"><strong>\u7da0\u8272\u80fd\u6e90\u3001\u6f01\u696d\u8cc7\u6e90\u3001\u6d77\u6d0b\u6c59\u67d3\u3001\u6c23\u5019\u8b8a\u9077\uff0c\u9019\u4e9b\u8a31\u591a\u4eba\u95dc\u5fc3\u7684\u8feb\u5207\u8b70\u984c\uff0c\u90fd\u8207\u6d77\u6d0b\u79d1\u5b78\u7814\u7a76\u606f\u606f\u76f8\u95dc\u3002\u53f0\u7063\u9019\u500b\u56db\u9762\u74b0\u6d77\u7684\u5cf6\u570b\uff0c\u5728\u6d77\u6d0b\u7814\u7a76\u7684\u9818\u57df\u4e2d\uff0c\u8d70\u904e\u4e86\u54ea\u4e9b\u7bf3\u8def\u85cd\u7e37\u7684\u6b77\u7a0b\uff0c\u672a\u4f86\u53c8\u6709\u4f55\u5c55\u671b\uff1f<\/strong><\/span><\/p>\n<p><strong>\u8a79\u68ee\uff5c\u81fa\u7063\u5927\u5b78\u6d77\u6d0b\u7814\u7a76\u6240\u6559\u6388<\/strong><\/p>\n<p>\u53f0\u7063\u53c3\u8207\u6d77\u6d0b\u7814\u7a76\uff0c\u8d77\u6b65\u5f97\u529b\u65bc\u7f8e\u570b\u7684\u5927\u529b\u5354\u52a9\u3002 1968 \u5e74\u53f0\u7063\u5927\u5b78\u6210\u7acb\u6d77\u6d0b\u7814\u7a76\u6240\uff0c\u662f\u7f8e\u570b\u9867\u554f\u5efa\u8b70\u4fc3\u6210\u7684\uff0c\u7b2c\u4e00\u8258\u6d77\u6d0b\u7814\u7a76\u8239\u300c\u4e5d\u9023\u8266\u300d\u4e5f\u662f\u7f8e\u570b\u6d77\u8ecd\u9000\u5f79\u8f49\u8d08\u3002\u7576\u6642\u570b\u5bb6\u9ad8\u5ea6\u91cd\u8996\u6d77\u6d0b\u7814\u7a76\uff0c\u526f\u7e3d\u7d71\u56b4\u5bb6\u6de6\u591a\u6b21\u8996\u5bdf\u53f0\u5927\u6d77\u7814\u6240\u8207\u4e5d\u9023\u8266\uff1b\u4e0d\u904e\u5be6\u969b\u4e0a\u5728\u4e0a\u500b\u4e16\u7d00\u7d50\u675f\u4e4b\u524d\uff0c\u9577\u9054\u4e09\u5341\u591a\u5e74\u7684\u671f\u9593\uff0c\u53f0\u7063\u50c5\u6709\u4e00\u4e9b\u96f6\u661f\u7684\u570b\u969b\u5408\u4f5c\u6d77\u6d0b\u7814\u7a76\u6848\u3002<\/p>\n<p>\u53f0\u7063\u7b2c\u4e00\u500b\u53c3\u8207\u7684\u570b\u969b\u5408\u4f5c\u6d77\u6d0b\u7814\u7a76\u6848\uff0c\u662f 1960 \u5e74\u4ee3\u5c31\u8457\u5730\u5229\u4e4b\u4fbf\uff0c\u53c3\u8207\u806f\u5408\u570b\u6559\u79d1\u6587\u7d44\u7e54\u4e3b\u5c0e\u7684\u300c\u806f\u5408\u63a2\u6e2c\u9ed1\u6f6e\u8a08\u756b\u300d (CSK) \uff1b\u4e0d\u904e\u96a8\u8457\u53f0\u7063\u9000\u51fa\u806f\u5408\u570b\uff0c\u672a\u518d\u53c3\u8207\u5f8c\u7e8c\u7814\u7a76\u8a08\u756b\uff0c\u5b78\u754c\u4e5f\u8ddf\u8457\u9677\u5165\u7814\u7a76\u4eba\u624d\u9752\u9ec3\u4e0d\u63a5\u7684\u7a98\u5883\u3002\u76f4\u5230 1980 \u5e74\u4ee3\uff0c\u53f0\u5927\u6d77\u6d0b\u6240\u57f9\u80b2\u51fa\u4f86\u7684\u7814\u7a76\u4eba\u624d\u9010\u6f38\u5145\u5be6\uff0c\u517c\u4e4b\u6d77\u7814\u4e00\u865f\u9069\u6642\u5b8c\u5de5\u4e0b\u6c34\uff0c\u4fc3\u6210\u300c\u9ed1\u6f6e\u908a\u7de3\u4ea4\u63db\u904e\u7a0b\u7814\u7a76\u8a08\u756b\u300d (KEEP) \uff0c\u6d77\u6d0b\u7814\u7a76\u7684\u5de8\u8f2a\u624d\u53c8\u958b\u59cb\u8f49\u52d5\u3002\u5230\u4e86 1990 \u5e74\u4ee3\uff0c\u6d77\u7814\u4e8c\u865f\u3001\u4e09\u865f\u5148\u5f8c\u670d\u5f79\uff0c\u53f0\u7063\u4e5f\u9806\u52e2\u52a0\u5165\u300c\u4e16\u754c\u6d77\u6d0b\u74b0\u6d41\u5be6\u9a57\u300d (WOCE) \u4ee5\u53ca\u300c\u5357\u6d77\u5b63\u98a8\u5be6\u9a57\u300d (SCSMEX) \u7b49\u570b\u969b\u5408\u4f5c\u8a08\u756b\uff0c\u6d77\u6d0b\u7814\u7a76\u9010\u6f38\u919e\u91c0\u80fd\u91cf\u3002<\/p>\n<p>\u53f0\u7063\u7684\u6d77\u6d0b\u7814\u7a76\u5728\u9032\u5165 21 \u4e16\u7d00\u4e4b\u5f8c\u84ec\u52c3\u767c\u5c55\uff0c\u4e0d\u4f46\u8207\u7f8e\u570b\u6d77\u8ecd\u7814\u7a76\u8655\u5171\u540c\u5408\u4f5c\u300c\u4e9e\u6d32\u6d77\u57df\u570b\u969b\u8072\u5b78\u5be6\u9a57\u300d (ASIAEX) \u3001\u300c\u5357\u6d77\u5167\u6ce2\u5be6\u9a57\u300d (IWISE) \u3001\u300c\u98b1\u98a8\u5be6\u9a57\u300d (ITOP) \u3001\u300c\u9ed1\u6f6e\u7de3\u8d77\u300d (OKMC) \u7b49\u8a08\u756b\uff0c\u81ea\u8eab\u4e5f\u958b\u59cb\u9032\u884c\u300c\u6771\u6d77\u9577\u671f\u89c0\u6e2c\u8207\u7814\u7a76\u300d (LORECS) \u3002\u96a8\u8457\u6d77\u7814\u4e94\u865f\u52a0\u5165\u7814\u7a76\u8239\u968a\uff0c\u53f0\u7063\u7684\u6d77\u6d0b\u7814\u7a76\u8db3\u8de1\u62d3\u5c55\u5230\u9060\u6d0b\u9031\u908a\u6d77\u57df\uff0c\u7814\u7a76\u7bc4\u7587\u4e5f\u56e0\u6b64\u66f4\u70ba\u5168\u9762\u3002<\/p>\n<p>\u53f0\u7063\u9019\u4e94\u5341\u5e74\u4f86\u7684\u6d77\u6d0b\u7814\u7a76\u6210\u679c\uff0c\u6fc3\u7e2e\u6210\u70ba\u79d1\u6280\u90e8\u7684\u300c\u6d77\u6d0b\u5b78\u9580\u8cc7\u6599\u5eab\u300d\u3002\u9019\u500b\u5b8c\u5168\u5c6c\u65bc\u6211\u5011\u81ea\u5df1\u7684\u6d77\u6d0b\u7814\u7a76\u8cc7\u6599\u5eab\uff0c\u5f59\u6574\u4e86\u5404\u8258\u6d77\u7814\u8239\u4ee5\u53ca\u6d77\u6d0b\u9328\u7887\u5100\u5668\u7684\u63a2\u6e2c\u7d50\u679c\uff0c\u9032\u884c\u7cfb\u7d71\u6027\u5206\u6790\u5f8c\uff0c\u505a\u6210\u6c34\u6587\u3001\u6d77\u6d41\u3001\u6c34\u6df1\u3001\u9707\u6e2c\u3001\u8072\u7d0d\u3001\u91cd\u529b\u3001\u5316\u5b78\u3001\u751f\u7269\u7b49\u7b49\u5404\u5f0f\u8cc7\u6599\uff0c\u4ee5\u4f9b\u7522\u696d\u754c\u8207\u5b78\u8853\u754c\u904b\u7528\u3002\u672a\u4f86\u96a8\u8457\u65b0\u4e00\u4ee3\u6d77\u6d0b\u7814\u7a76\u8239\u52a0\u5165\u9060\u6d0b\u6d77\u7814\u884c\u5217\uff0c\u7b2c\u4e8c\u3001\u7b2c\u4e09\u4ee3\u7684\u6d77\u7814\u4eba\u624d\u63a5\u529b\u6295\u5165\u4e4b\u4e0b\uff0c\u53f0\u7063\u7684\u6d77\u6d0b\u7814\u7a76\u5c07\u7522\u751f\u65b0\u601d\u7dad\uff0c\u62d3\u5c55\u51fa\u524d\u6240\u672a\u6709\u7684\u53ef\u80fd\u6027\u3002<\/p>\n","protected":false},"excerpt":{"rendered":"<p>\u6d77\u6d0b\u5fae\u751f\u7269\u7a2e\u985e\u7e41\u591a\uff0c\u5305\u62ec\u81ea\u71df\u7d30\u83cc\u3001\u7570\u71df\u7d30\u83cc\u3001\u53e4\u7d30\u83cc\u3001\u75c5\u6bd2\u3001\u7e96\u6bdb\u87f2\u3001\u97ad\u6bdb\u87f2\u3001\u771f\u83cc\u3001\u85fb\u985e\u7b49\u7b49\uff0c\u6700\u65e9\u53ef\u8ffd\u6eaf\u5230 35 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