{"id":34176,"date":"2019-07-15T00:00:28","date_gmt":"2019-07-14T16:00:28","guid":{"rendered":"https:\/\/case.ntu.edu.tw\/blog\/?p=34176"},"modified":"2023-06-06T11:18:00","modified_gmt":"2023-06-06T03:18:00","slug":"%e4%bd%a0%e7%9a%84%e7%99%bd%e5%a4%a9%e8%ba%ab%e9%ab%94%e4%b8%8d%e6%98%af%e4%bd%a0%e7%9a%84%e6%99%9a%e4%b8%8a%e8%ba%ab%e9%ab%94-%e7%94%9f%e7%90%86%e6%99%82%e9%90%98%e5%a6%82%e4%bd%95%e7%9c%8b","status":"publish","type":"post","link":"https:\/\/case.ntu.edu.tw\/blog\/?p=34176","title":{"rendered":"\u3010\u79d1\u5b78\u53f2\u6c99\u9f8d\u3011\u3008\u4f60\u7684\u767d\u5929\u8eab\u9ad4\u4e0d\u662f\u4f60\u7684\u665a\u4e0a\u8eab\u9ad4\u2014\u2014\u751f\u7406\u6642\u9418\u5982\u4f55\u770b\u898b\u85cd\u5149\u3009&#038;\u3008\u5927\u8166\u4e2d\u7684\u85cd\u8272\u7cbe\u9748\u3009"},"content":{"rendered":"<p style=\"text-align: center;\"><iframe src=\"https:\/\/www.youtube.com\/embed\/IdfoDX2r0Qo\" width=\"560\" height=\"315\" frameborder=\"0\" allowfullscreen=\"allowfullscreen\"><\/iframe><\/p>\n<p><span style=\"color: #993300;\"><strong>\u3008\u4f60\u7684\u767d\u5929\u8eab\u9ad4\u4e0d\u662f\u4f60\u7684\u665a\u4e0a\u8eab\u9ad4\u2014\u2014\u751f\u7406\u6642\u9418\u5982\u4f55\u770b\u898b\u85cd\u5149\u3009<\/strong><\/span><\/p>\n<p><span style=\"color: #800080;\"><strong>\u5730\u7403\u6bcf 24 \u5c0f\u6642\u81ea\u8f49\u4e00\u5468\uff0c\u9019\u500b\u898f\u5f8b\u7684\u65e5\u591c\u9031\u671f\uff0c\u5c0d\u65bc\u5730\u7403\u4e0a\u7684\u751f\u7269\u9020\u6210\u6975\u986f\u8457\u7684\u5f71\u97ff\u3002\u70ba\u4e86\u9069\u61c9\u9019\u500b\u898f\u5f8b\u7684\u9031\u671f\u8b8a\u5316\uff0c\u751f\u7269\u9ad4\u5167\u5f62\u6210\u4e86\u4e00\u500b\u5167\u751f\u6027\u7684\u751f\u7406\u6642\u9418\uff0c\u5e6b\u52a9\u500b\u9ad4\u6e96\u5099\u5373\u5c07\u4f86\u81e8\u7684\u5916\u5728\u8b8a\u5316\u3002\u672c\u8b1b\u6b21\u8aaa\u660e\u751f\u7269\u9ad4\u900f\u904e\u4f55\u7a2e\u6a5f\u5236\u8abf\u7bc0\u751f\u7406\u6642\u9418\uff0c\u4ee5\u53ca\u4eba\u5011\u5728\u73fe\u4ee3\u793e\u6703\u4e2d\u8a72\u6ce8\u610f\u4e9b\u4ec0\u9ebc\u4e8b\u60c5\uff0c\u907f\u514d\u751f\u7406\u6642\u9418\u53d7\u5230\u5e72\u64fe\u3002<\/strong><\/span><\/p>\n<p><strong>\u8b1b\u5e2b\uff5c\u81fa\u7063\u5927\u5b78\u751f\u547d\u79d1\u5b78\u7cfb\u6559\u6388 \u9673\u793a\u570b<br \/>\n\u6574\u7406\u64b0\u6587\uff5c\u9ad8\u82f1\u54f2<br \/>\n<\/strong><\/p>\n<p>\u5730\u7403\u65e5\u591c\u9031\u671f\u7684\u5dee\u7570\uff0c\u6700\u660e\u986f\u7684\u5730\u65b9\u5728\u65bc\u5149\u7167\u5ea6\uff0c\u767d\u5929\u8ddf\u665a\u4e0a\u7684\u5149\u91cf\u76f8\u5dee\u4e00\u5104\u500d\u4ee5\u4e0a\uff1b\u5176\u6b21\u662f\u6c23\u6eab\u5dee\u7570\uff0c\u5373\u4f7f\u5728\u8b8a\u5316\u6bd4\u8f03\u6eab\u548c\u7684\u5340\u57df\uff0c\u65e5\u591c\u6eab\u5dee\u4e5f\u6703\u9054\u5230\u651d\u6c0f\u5341\u5ea6\u4ee5\u4e0a\u3002\u751f\u7269\u70ba\u4e86\u9069\u61c9\u9019\u500b\u65e2\u660e\u986f\u53c8\u898f\u5f8b\u7684\u8b8a\u5316\uff0c\u9010\u6f38\u884d\u751f\u51fa\u4e00\u5957\u8abf\u7bc0\u8eab\u9ad4\u6a5f\u80fd\u7684\u751f\u7406\u6642\u9418\u3002\u751f\u7406\u6642\u9418\u7684\u82f1\u6587 circadian clock \uff0c\u6b63\u662f\u300c\u5927\u7d04\u300d (circa) \u8ddf\u300c\u4e00\u65e5\u300d (dian) \u9019\u5169\u500b\u5e0c\u81d8\u5b57\u7d44\u5408\u800c\u6210\uff1b\u8209\u51e1\u9ad4\u5167\u7684\u892a\u9ed1\u6fc0\u7d20\u3001\u6838\u5fc3\u9ad4\u6eab\u3001\u4e09\u9178\u7518\u6cb9\u8102\uff0c\u4ee5\u81f3\u65bc\u4e3b\u89c0\u8b66\u89ba\u6027\u3001\u53cd\u61c9\u6642\u9593\u7b49\u7b49\uff0c\u90fd\u6703\u96a8\u8457\u751f\u7406\u6642\u9418\u800c\u51fa\u73fe\u898f\u5f8b\u6027\u7684\u8b8a\u7570\u3002<\/p>\n<p>\u96d6\u7136\u81ea\u53e4\u4ee5\u4f86\u5c31\u6709\u4eba\u6ce8\u610f\u5230\u751f\u7406\u6642\u9418\u9019\u500b\u73fe\u8c61\uff0c\u7136\u800c\u4e00\u76f4\u8981\u5230 1950 \u5e74\u4ee3\uff0c\u79d1\u5b78\u754c\u624d\u958b\u59cb\u5c0d\u9019\u500b\u73fe\u8c61\u6709\u7cfb\u7d71\u6027\u7684\u7814\u7a76\u3002\u5fb7\u570b\u751f\u7406\u5b78\u5bb6\u827e\u8a31\u592b (J\u00fcrgen Aschoff) \u4ee5\u53ca\u7f8e\u570b\u751f\u7269\u5b78\u5bb6\u76ae\u767b\u5353 (Colin Pittendrigh) \uff0c\u662f\u5f15\u9818\u9019\u500b\u7814\u7a76\u9818\u57df\u7684\u5148\u884c\u8005\uff0c\u4ed6\u5011\u79c1\u4ea4\u751a\u7be4\uff0c\u5c0d\u65bc\u5982\u4f55\u8abf\u7bc0\u751f\u7406\u6642\u9418\uff0c\u537b\u6709\u8457\u622a\u7136\u4e0d\u540c\u7684\u898b\u89e3\uff1a\u827e\u8a31\u592b\u4e3b\u5f35\u8abf\u7bc0\u5149\u7167\u5ea6\uff0c\u76ae\u767b\u5353\u537b\u8a8d\u70ba\u9078\u5c0d\u6642\u9593\u624d\u662f\u91cd\u9ede\u3002<\/p>\n<p>\u827e\u8a31\u592b\u5229\u7528\u4e8c\u6230\u7559\u4e0b\u7684\u5730\u4e0b\u7889\u5821\u505a\u5be6\u9a57\uff0c\u8b93\u53d7\u6e2c\u8005\u5728\u4e0d\u898b\u5929\u65e5\u7684\u74b0\u5883\u4e2d\u751f\u6d3b\uff0c\u4e26\u7d00\u9304\u4ed6\u5011\u7684\u7761\u7720\u6642\u9593\uff0c\u767c\u73fe\u53d7\u6e2c\u8005\u7684\u7761\u7720\u6642\u6bb5\uff0c\u5f88\u898f\u5f8b\u5730\u6bcf\u5929\u63a8\u9072\u5927\u7d04\u534a\u500b\u5c0f\u6642\uff0c\u4e5f\u5c31\u662f\u751f\u7406\u6642\u9418\u4ee5 24.5 \u500b\u5c0f\u6642\u70ba\u4e00\u500b\u9031\u671f\u2014\u2014\u9019\u5c31\u662f\u70ba\u4ec0\u9ebc\u751f\u7406\u6642\u9418\u7684\u82f1\u6587\u5b57\u662f\u300c\u5927\u7d04\u4e00\u65e5\u300d\u3002\u9019\u610f\u5473\u8457\u751f\u7406\u6642\u9418\u4e26\u4e0d\u662f\u5b8c\u5168\u5207\u5408\u5730\u7403\u81ea\u8f49\u7684\u898f\u5f8b\uff0c\u800c\u4e14\u5f8c\u7e8c\u7814\u7a76\u9084\u767c\u73fe\u6bcf\u500b\u7269\u7a2e\u7684\u751f\u7406\u6642\u9418\u4e5f\u6709\u4e9b\u8a31\u5dee\u7570\uff0c\u56e0\u6b64\u751f\u7269\u5fc5\u9808\u4f9d\u64da\u81ea\u7136\u74b0\u5883\u4e2d\u5149\u7167\u5ea6\u7684\u8b8a\u5316\uff0c\u8abf\u7bc0\u81ea\u5df1\u7684\u751f\u7406\u6642\u9418\u3002<\/p>\n<p>\u4eba\u985e\u900f\u904e\u773c\u775b\u7684\u611f\u5149\u7d30\u80de\uff0c\u611f\u61c9\u5149\u7167\u5ea6\u7684\u8b8a\u5316\u3002\u7136\u800c\u5947\u602a\u7684\u662f\uff0c\u6709\u4e9b\u611f\u5149\u7d30\u80de\u56e0\u75c5\u6216\u56e0\u50b7\u840e\u7e2e\u7684\u4eba\uff0c\u4ed6\u5011\u7684\u8eab\u9ad4\u7167\u6a23\u53ef\u4ee5\u8abf\u6574\u6642\u5dee\uff0c\u5f8c\u4f86\u767c\u73fe\u662f\u4e00\u7a2e\u53eb\u505a\u8996\u9ed1\u8cea (melanopsin) \u7684\u611f\u5149\u86cb\u767d\uff0c\u4e0d\u9700\u8981\u7d93\u7531\u8996\u9310\u4ee5\u53ca\u8996\u611f\u7d30\u80de\uff0c\u4e5f\u53ef\u4ee5\u611f\u61c9\u5149\u7167\u8b8a\u5316\uff0c\u4e26\u4e14\u628a\u8a0a\u606f\u50b3\u905e\u5230\u8ca0\u8cac\u8abf\u7bc0\u751f\u7406\u6642\u9418\u7684\u8996\u4ea4\u53c9\u4e0a\u6838\u3002\u7531\u65bc\u8996\u9ed1\u8cea\u6240\u80fd\u611f\u61c9\u5230\u7684\u5149\u6ce2\u9577\uff0c\u6b63\u597d\u662f\u843d\u5728\u85cd\u5149\u7684\u5340\u6bb5\uff0c\u5982\u679c\u8996\u9ed1\u8cea\u5728\u591c\u9593\u63a5\u6536\u5230\u904e\u591a\u85cd\u5149\uff0c\u8996\u4ea4\u53c9\u4e0a\u6838\u4fbf\u6703\u4ee5\u70ba\u73fe\u5728\u662f\u6709\u5929\u7a7a\u85cd\u7684\u767d\u5929\uff0c\u628a\u8eab\u9ad4\u8abf\u7bc0\u6210\u9069\u61c9\u767d\u5929\u6d3b\u52d5\u7684\u72c0\u614b\uff0c\u9020\u6210\u751f\u7406\u6a5f\u80fd\u7684\u932f\u4e82\u3002<\/p>\n<p>\u4e86\u89e3\u4eba\u9ad4\u8abf\u7bc0\u751f\u7406\u6642\u9418\u7684\u6a5f\u5236\u5f8c\uff0c\u6211\u5011\u5c31\u80fd\u63a1\u53d6\u76f8\u5c0d\u61c9\u7684\u7b56\u7565\u2014\u2014\u76e1\u53ef\u80fd\u8b93\u751f\u6d3b\u4f5c\u606f\u5728\u4e00\u6bb5\u6642\u9593\u5167\u4fdd\u6301\u56fa\u5b9a\uff08\u56fa\u5b9a\u767d\u73ed\u6216\u56fa\u5b9a\u591c\u73ed\uff09\uff0c\u4ee5\u53ca\u5728\u61c9\u8a72\u4f11\u606f\u6642\u907f\u514d\u66b4\u9732\u5728\u85cd\u5149\u74b0\u5883\u4e4b\u4e2d\uff08\u6bd4\u65b9\u8aaa\u4f7f\u7528\u96fb\u5b50\u7522\u54c1\uff09\u2014\u2014\u8b93\u81ea\u5df1\u7684\u751f\u7406\u6642\u9418\u66f4\u52a0\u7a69\u5b9a\u3002<\/p>\n<p>&nbsp;<\/p>\n<p><span style=\"color: #993300;\"><strong>\u3008\u5927\u8166\u4e2d\u7684\u85cd\u8272\u7cbe\u9748\u3009<\/strong><\/span><\/p>\n<p><span style=\"color: #800080;\"><strong>\u52d5\u7269\u5c0d\u65bc\u4ee3\u8868\u300c\u665a\u9910\u300d\u7684\u9234\u8072\uff0c\u9664\u975e\u8655\u5728\u6beb\u7121\u8b66\u89ba\u7684\u7761\u7720\uff0c\u4e0d\u7136\u5728\u4e00\u822c\u60c5\u5f62\u4e0b\uff0c\u4e0d\u53ef\u80fd\u6703\u5ffd\u7565\u9019\u9ebc\u91cd\u8981\u7684\u8a0a\u606f\uff1b\u7136\u800c\u7576\u7260\u5011\u8655\u65bc\u9700\u8981\u4fdd\u6301\u9ad8\u5ea6\u8b66\u89ba\uff0c\u7372\u53d6\u5229\u76ca\u9060\u5927\u65bc\u4e00\u9910\u7684\u60c5\u5f62\u6642\uff0c\u5927\u8166\u5fc5\u9808\u8981\u5ffd\u7565\u9234\u8072\u7684\u5e72\u64fe\uff0c\u4ee5\u4fbf\u5c08\u6ce8\u65bc\u8655\u7406\u773c\u4e0b\u6240\u7372\u5f97\u7684\u8a0a\u606f\u3002\u5927\u8166\u4e2d\u8ca0\u8cac\u8655\u7406\u9019\u7a2e\u5f48\u6027\u9078\u64c7\u7684\uff0c\u662f\u4e00\u500b\u7a31\u70ba\u300c\u85cd\u6591\u6838\u300d\u7684\u5c0f\u7d50\u69cb\uff0c\u672c\u8b1b\u6b21\u4ecb\u7d39\u9019\u500b\u6700\u65e9\u53d7\u5230\u7814\u7a76\u7684\u5927\u8166\u5340\u57df\u4e4b\u4e00\uff0c\u5176\u904b\u4f5c\u539f\u7406\u53ca\u91cd\u8981\u6027\u3002<\/strong><\/span><\/p>\n<p><strong>\u8b1b\u5e2b\uff5c\u81fa\u7063\u5927\u5b78\u751f\u547d\u79d1\u5b78\u7cfb\u6559\u6388 \u9594\u660e\u6e90<\/strong><br \/>\n<strong>\u6574\u7406\u64b0\u6587\uff5c\u9ad8\u82f1\u54f2<\/strong><\/p>\n<p>\u52d5\u7269\u5c0d\u65bc\u6240\u8655\u74b0\u5883\u7684\u53cd\u61c9\uff0c\u5927\u62b5\u5206\u70ba\u5169\u7a2e\uff1a\u4e00\u662f\u5c0d\u65bc\u5df2\u77e5\u7684\u734e\u52f5\u4f86\u6e90\u9032\u884c\u300c\u5229\u7528\u300d\uff0c\u4e8c\u662f\u5c0d\u65bc\u672a\u77e5\u7684\u74b0\u5883\u9032\u884c\u300c\u63a2\u7d22\u300d\uff0c\u4ee5\u7372\u53d6\u6bd4\u5df2\u77e5\u734e\u52f5\u66f4\u8c50\u5bcc\u3001\u66f4\u7a69\u5b9a\u7684\u4f86\u6e90\u3002\u9019\u5169\u7a2e\u53cd\u61c9\u4e4b\u9593\u6709\u5176\u4ea4\u63db\u4ee3\u50f9\uff1a\u4e00\u500b\u53ea\u6eff\u8db3\u65bc\u300c\u5229\u7528\u300d\u7684\u500b\u9ad4\uff0c\u96d6\u7136\u53ef\u4ee5\u7a69\u5b9a\u5730\u7372\u53d6\u56fa\u5b9a\u734e\u52f5\uff0c\u537b\u653e\u68c4\u4e86\u9032\u4e00\u6b65\u6539\u5584\u751f\u6d3b\u7684\u6a5f\u6703\uff1b\u800c\u4e00\u500b\u52c7\u65bc\u9032\u884c\u300c\u63a2\u7d22\u300d\u7684\u500b\u9ad4\uff0c\u96d6\u7136\u6709\u6a5f\u6703\u6539\u5584\u5176\u751f\u6d3b\u689d\u4ef6\uff0c\u537b\u4e5f\u5fc5\u9808\u627f\u64d4\u5f92\u52de\u7121\u529f\uff0c\u6d6a\u8cbb\u80fd\u91cf\u7684\u98a8\u96aa\u3002<\/p>\n<p>\u8ca0\u8cac\u8655\u7406\u52d5\u7269\u63a1\u53d6\u300c\u5229\u7528\u300d\u6216\u300c\u63a2\u7d22\u300d\u884c\u70ba\u7684\u5927\u8166\u5340\u57df\uff0c\u662f\u4f4d\u65bc\u4e2d\u8166\u7684\u85cd\u6591\u6838 (locus coeruleus) \u3002\u85cd\u6591\u6838\u9867\u540d\u601d\u7fa9\u5448\u73fe\u85cd\u8272\uff0c\u9019\u662f\u56e0\u70ba\u5b83\u91cb\u653e\u51fa\u4f86\uff0c\u4e00\u7a2e\u53eb\u505a\u6b63\u814e\u4e0a\u817a\u7d20 (norepinephrine) \u7684\u795e\u7d93\u50b3\u5c0e\u7269\u8cea\uff0c\u672c\u8eab\u5c31\u6703\u5f62\u6210\u9ed1\u8272\u7d20\uff0c\u5982\u679c\u6c92\u6709\u7d2f\u7a4d\u5f88\u591a\u7684\u8a71\uff0c\u770b\u8d77\u4f86\u5c31\u6703\u63a5\u8fd1\u85cd\u9ed1\u8272\u3002\u85cd\u6591\u6838\u662f\u5927\u8166\u4e2d\u6b63\u814e\u4e0a\u817a\u7d20\u7684\u4e3b\u8981\u4f9b\u61c9\u8005\uff0c\u5e7e\u4e4e\u6240\u6709\u7684\u5927\u8166\u5340\u57df\uff0c\u90fd\u542b\u6709\u7dfb\u5bc6\u7684\u6b63\u814e\u4e0a\u817a\u7d20\u795e\u7d93\u7e96\u7dad\u3002\u85cd\u6591\u6838\u5177\u6709\u5168\u9762\u6027\u7684\u884c\u70ba\u8abf\u7bc0\u529f\u80fd\uff0c\u5b83\u900f\u904e\u6b63\u814e\u4e0a\u817a\u7d20\u7cfb\u7d71\uff0c\u8abf\u7bc0\u5927\u8166\u7684\u7761\u7720\u8207\u6e05\u9192\u9031\u671f\uff0c\u81ea\u4e3b\u795e\u7d93\u529f\u80fd\u63a7\u5236\uff0c\u60c5\u7dd2\u72c0\u614b\u8abf\u7bc0\uff0c\u5b78\u7fd2\u8207\u8a18\u61b6\uff0c\u4ee5\u53ca\u75db\u89ba\u7b49\u7b49\u3002<\/p>\n<p>\u7814\u7a76\u8005\u85c9\u7531\u63a7\u5236\u734e\u52f5\u4ee5\u53ca\u6e2c\u91cf\u5927\u8166\u653e\u96fb\u7b49\u7b49\u5be6\u9a57\u624b\u6bb5\uff0c\u4e86\u89e3\u85cd\u6591\u6838\u5c0d\u65bc\u8eab\u9ad4\u6a5f\u80fd\u7684\u8abf\u7bc0\u6a5f\u5236\uff0c\u4ee5\u53ca\u5927\u8166\u63a5\u6536\u523a\u6fc0\uff0c\u9032\u884c\u6c7a\u7b56\uff0c\u505a\u51fa\u53cd\u61c9\u7b49\u7b49\uff0c\u4e00\u7cfb\u5217\u50b3\u5c0e\u904e\u7a0b\u7684\u653e\u96fb\u500d\u7387\u503c\u3002\u5c0d\u85cd\u6591\u6838\u7684\u8abf\u7bc0\u6a5f\u5236\u6709\u4e00\u5b9a\u4e86\u89e3\u4e4b\u5f8c\uff0c\u5c31\u53ef\u900f\u904e\u523a\u6fc0\u85cd\u6591\u6838\u55ae\u795e\u7d93\u5143\u653e\u96fb\u6d3b\u52d5\u7b49\u7b49\u624b\u6bb5\uff0c\u9032\u884c\u4e3b\u52d5\u7684\u8abf\u7bc0\u63a7\u7ba1\uff0c\u6bd4\u65b9\u8aaa\u8981\u63d0\u9ad8\u63a2\u7d22\u74b0\u5883\u654f\u92b3\u5ea6\u6642\u523a\u6fc0\u653e\u96fb\uff0c\u9700\u8981\u5c08\u6ce8\u8655\u7406\u773c\u4e0b\u4e8b\u52d9\u6642\u5c31\u6291\u5236\u653e\u96fb\u3002\u5176\u4ed6\u8af8\u5982\u589e\u5f37\u8a8d\u77e5\u529f\u80fd\uff0c\u63d0\u9ad8\u52d5\u6a5f\u6c34\u5e73\uff0c\u4fc3\u9032\u4ea4\u611f\u795e\u7d93\u6d3b\u52d5\u7b49\u7b49\uff0c\u4e5f\u90fd\u662f\u85cd\u6591\u6838\u5728\u81e8\u5e8a\u4e0a\u53ef\u80fd\u7684\u76f8\u95dc\u61c9\u7528\uff0c\u53ef\u5354\u52a9\u60a3\u8005\u64fa\u812b\u751f\u6d3b\u4e0a\u4e0d\u5fc5\u8981\u7684\u5e72\u64fe\u3002<\/p>\n<p>&nbsp;<\/p>\n<p><a href=\"https:\/\/line.me\/R\/ti\/p\/%40fcx6182w\" target=\"_blank\" rel=\"noopener noreferrer\"><img decoding=\"async\" class=\"alignnone\" src=\"https:\/\/scdn.line-apps.com\/n\/line_add_friends\/btn\/zh-Hant.png\" alt=\"\u52a0\u5165\u597d\u53cb\" width=\"212\" height=\"36\" border=\"0\"><\/a><\/p>\n","protected":false},"excerpt":{"rendered":"<p>\u5730\u7403\u6bcf 24 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