{"id":20895,"date":"2015-03-03T00:05:49","date_gmt":"2015-03-02T16:05:49","guid":{"rendered":"https:\/\/case.ntu.edu.tw\/blog\/?p=20895"},"modified":"2018-07-13T00:25:13","modified_gmt":"2018-07-12T16:25:13","slug":"%e3%80%90%e5%9c%8b%e9%9a%9b%e5%9c%9f%e5%a3%a4%e5%b9%b4%e5%b0%88%e9%a1%8c%e3%80%91%e6%a0%b9%e7%98%a4%e8%8f%8crhizobium%e8%88%87%e4%ba%ba","status":"publish","type":"post","link":"https:\/\/case.ntu.edu.tw\/blog\/?p=20895","title":{"rendered":"\u3010\u570b\u969b\u571f\u58e4\u5e74\u5c08\u984c\u3011\u6839\u7624\u83cc(Rhizobium)\u8207\u4eba"},"content":{"rendered":"<figure id=\"attachment_20664\" aria-describedby=\"caption-attachment-20664\" style=\"width: 256px\" class=\"wp-caption alignleft\"><img fetchpriority=\"high\" decoding=\"async\" class=\"wp-image-20664\" src=\"http:\/\/case.ntu.edu.tw\/blog\/wp-content\/uploads\/2015\/02\/Soybean.USDA_.jpg\" alt=\"\u5927\u8c46\u3002\u5716\u7247\u4f86\u6e90\uff1awiki\" width=\"256\" height=\"380\" srcset=\"https:\/\/case.ntu.edu.tw\/blog\/wp-content\/uploads\/2015\/02\/Soybean.USDA_.jpg 640w, https:\/\/case.ntu.edu.tw\/blog\/wp-content\/uploads\/2015\/02\/Soybean.USDA_-600x892.jpg 600w\" sizes=\"(max-width: 256px) 100vw, 256px\" \/><figcaption id=\"caption-attachment-20664\" class=\"wp-caption-text\">\u5927\u8c46\u3002\u5716\u7247\u4f86\u6e90\uff1awiki<\/figcaption><\/figure>\n<p><strong>\u64b0\u6587\uff5c<\/strong>\u8449\u7da0\u8212<\/p>\n<p>\u81ea\u5f9e\u4eba\u985e\u5728\u4e00\u842c\u4e00\u5343\u5e74\u524d\u5b78\u6703\u4e86\u8fb2\u8015\u4ee5\u5f8c\uff0c\u5982\u4f55\u63d0\u5347\u8fb2\u4f5c\u7269\u7684\u7522\u91cf\u4fbf\u6210\u70ba\u4eba\u5011\u5ff5\u8332\u5728\u8332\u7684\u8b70\u984c\u3002\u4f7f\u7528\u52d5\u7269\u7684\u6392\u6cc4\u7269\u3001\u4ee5\u53ca\u6536\u96c6\u690d\u7269\u7684\u843d\u8449\u3001\u5bb6\u5ead\u5783\u573e\u88fd\u4f5c\u5806\u80a5\uff0c\u9019\u4e9b\u65b9\u5f0f\u90fd\u5728\u8a66\u9a57\u4e2d\u767c\u73fe\u53ef\u4ee5\u63d0\u5347\u7522\u91cf\uff1b\u800c\u53ef\u80fd\u65e9\u5728\u53e4\u57c3\u53ca\u6642\u4ee3\uff0c\u4eba\u5011\u4fbf\u5df2\u7d93\u767c\u73fe\u7a2e\u690d\u8c46\u79d1\u690d\u7269(legumes)\u5982\u5927\u8c46\u3001\u8c4c\u8c46\u7b49\uff0c\u53ef\u4ee5\u4f7f\u571f\u58e4\u80a5\u6c83\u3002<\/p>\n<p>\u4e2d\u570b\u5728\u6f22\u671d\u6642\u5df2\u61c2\u5f97\u5229\u7528\u5927\u8c46\u8207\u5c0f\u7c73\u3001\u5c0f\u9ea5\u8f2a\u4f5c(\u53c31)\uff0c\u7dad\u6301\u571f\u58e4\u7684\u80a5\u529b\uff1b\u540c\u6a23\u7684\u6982\u5ff5\uff0c\u5728\u6b50\u6d32\u5927\u7d04\u4e00\u5343\u5e74\u5f8c\u624d\u51fa\u73fe\u3002\u4f46\u662f\uff0c\u4e0d\u8ad6\u662f\u57c3\u53ca\u3001\u4e2d\u570b\u3001\u6216\u662f\u6b50\u6d32\uff0c\u5c0d\u65bc\u70ba\u4f55\u8c46\u79d1\u690d\u7269\u53ef\u4ee5\u63d0\u5347\u571f\u58e4\u7684\u80a5\u529b\u9019\u4ef6\u4e8b\uff0c\u90fd\u662f\u77e5\u5176\u7136\u800c\u4e0d\u77e5\u5176\u6240\u4ee5\u7136\u3002<\/p>\n<p>\u9019\u500b\u8b0e\u984c\uff0c\u76f4\u5230\u5341\u4e5d\u4e16\u7d00\u672b\u624d\u88ab\u89e3\u958b\u30021888\u5e74\uff0c\u5fb7\u570b\u7684\u8fb2\u696d\u5316\u5b78\u5bb6Hermann Hellriegel(\u4e0b\u5716)\u8207 Hermann Wilfarth\u767c\u73fe\u6839\u7624\u53ef\u4ee5\u5c07\u7a7a\u6c23\u4e2d\u7684\u6c2e(nitrogen)\u8f49\u8b8a\u70ba\u6c28(ammonia)\u3002\u5728\u540c\u4e00\u5e74\uff0c\u6839\u7624\u83cc\u4e5f\u88ab\u8377\u862d\u7684Martinus W. Beijerinck(\u4e0b\u5716)\u767c\u73fe\u4e86(\u53c32)\u3002<!--more--><\/p>\n<figure id=\"attachment_20666\" aria-describedby=\"caption-attachment-20666\" style=\"width: 284px\" class=\"wp-caption aligncenter\"><img decoding=\"async\" class=\"wp-image-20666\" src=\"http:\/\/case.ntu.edu.tw\/blog\/wp-content\/uploads\/2015\/02\/Hermann_Hellriegel.jpg\" alt=\"Hermann Hellriegel. \u5716\u7247\u4f86\u6e90\uff1awiki\" width=\"284\" height=\"400\" srcset=\"https:\/\/case.ntu.edu.tw\/blog\/wp-content\/uploads\/2015\/02\/Hermann_Hellriegel.jpg 463w, https:\/\/case.ntu.edu.tw\/blog\/wp-content\/uploads\/2015\/02\/Hermann_Hellriegel-213x300.jpg 213w\" sizes=\"(max-width: 284px) 100vw, 284px\" \/><figcaption id=\"caption-attachment-20666\" class=\"wp-caption-text\">Hermann Hellriegel. \u5716\u7247\u4f86\u6e90\uff1awiki<\/figcaption><\/figure>\n<figure id=\"attachment_20667\" aria-describedby=\"caption-attachment-20667\" style=\"width: 284px\" class=\"wp-caption aligncenter\"><img decoding=\"async\" class=\"wp-image-20667\" src=\"http:\/\/case.ntu.edu.tw\/blog\/wp-content\/uploads\/2015\/02\/Martinus_Willem_Beijerinck.png\" alt=\"Martinus W. Beijerinck.\u5716\u7247\u4f86\u6e90\uff1awiki\" width=\"284\" height=\"384\" srcset=\"https:\/\/case.ntu.edu.tw\/blog\/wp-content\/uploads\/2015\/02\/Martinus_Willem_Beijerinck.png 640w, https:\/\/case.ntu.edu.tw\/blog\/wp-content\/uploads\/2015\/02\/Martinus_Willem_Beijerinck-221x300.png 221w\" sizes=\"(max-width: 284px) 100vw, 284px\" \/><figcaption id=\"caption-attachment-20667\" class=\"wp-caption-text\">Martinus W. Beijerinck.\u5716\u7247\u4f86\u6e90\uff1awiki<\/figcaption><\/figure>\n<p>\u96d6\u7136\u5728\u904e\u53bb\u6839\u7624\u83cc\u66fe\u7d93\u4e00\u5ea6\u88ab\u8aa4\u6703\u70ba\u771f\u83cc(\u4e5f\u771f\u7684\u6709\u771f\u83cc\u8207\u690d\u7269\u5171\u751f\uff0c\u4e0d\u904e\u5171\u751f\u7684\u65b9\u5f0f\u8207\u6839\u7624\u83cc\u4e0d\u540c)\uff0c\u7576\u6642Beijerinck\u5c07\u6839\u7624\u83cc\u547d\u540d\u70ba<em>Bacillus radicicola<\/em>\uff0c\u5f8c\u4f86\u624d\u88ab\u91cd\u65b0\u5206\u985e\u5230<em>Rhizobium<\/em>\u9019\u4e00\u5c6c\u3002<\/p>\n<p>\u5f9e\u90a3\u6642\u5019\u958b\u59cb\uff0c\u6839\u7624\u83cc\u5c31\u6210\u70ba\u7814\u7a76\u8005\u7684\u8ab2\u984c\u4e4b\u4e00\u3002\u8c4c\u8c46\u6839\u7624\u83cc(<em>Rhizobium leguminosarum)<\/em>\u7684\u57fa\u56e0\u9ad4\u5df2\u7d93\u57282006\u5e74\u5b9a\u5e8f\u5b8c\u6210(\u5982\u4e0b\u5716)\uff0c\u9664\u4e86\u5b83\u672c\u8eab\u7684\u67d3\u8272\u9ad4(\u53f3\u4e0a)\u4ee5\u5916\uff0c\u6839\u7624\u83cc\u9084\u6709\u516d\u500b\u8cea\u9ad4\uff0c\u5176\u4e2d\u8207\u690d\u7269\u5171\u751f\u76f8\u95dc\u7684<em>nod(<\/em>\u6839\u7624\u5f62\u6210\uff0cnodulation of the host plant)\u4ee5\u53ca<em>nit<\/em>\u8207<em>fix(<\/em>\u56fa\u6c2e)\uff0c\u90fd\u4f4d\u65bc\u7b2c\u4e09\u5927\u7684\u8cea\u9ad4pRL10\u4e0a\u3002<\/p>\n<figure id=\"attachment_20668\" aria-describedby=\"caption-attachment-20668\" style=\"width: 450px\" class=\"wp-caption aligncenter\"><a href=\"http:\/\/case.ntu.edu.tw\/blog\/wp-content\/uploads\/2015\/02\/gb-2006-7-4-r34-1.jpg\"><img loading=\"lazy\" decoding=\"async\" class=\"wp-image-20668\" src=\"http:\/\/case.ntu.edu.tw\/blog\/wp-content\/uploads\/2015\/02\/gb-2006-7-4-r34-1.jpg\" alt=\"\u8c4c\u8c46\u6839\u7624\u83cc\u7684\u57fa\u56e0\u9ad4\u3002\u5716\u7247\u4f86\u6e90\uff1aGenome Biology\" width=\"450\" height=\"433\" srcset=\"https:\/\/case.ntu.edu.tw\/blog\/wp-content\/uploads\/2015\/02\/gb-2006-7-4-r34-1.jpg 600w, https:\/\/case.ntu.edu.tw\/blog\/wp-content\/uploads\/2015\/02\/gb-2006-7-4-r34-1-300x288.jpg 300w\" sizes=\"(max-width: 450px) 100vw, 450px\" \/><\/a><figcaption id=\"caption-attachment-20668\" class=\"wp-caption-text\">\u8c4c\u8c46\u6839\u7624\u83cc\u7684\u57fa\u56e0\u9ad4\u3002\u5716\u7247\u4f86\u6e90\uff1aGenome Biology<\/figcaption><\/figure>\n<p>\u6839\u7624\u83cc\u8207\u690d\u7269\u4e4b\u9593\u7684\u4e92\u52d5\u662f\u76f8\u7576\u8907\u96dc\u7684\u3002\u6839\u7624\u83cc\u539f\u672c\u662f\u571f\u58e4\u4e2d\u81ea\u7531\u751f\u6d3b\u7684\u683c\u862d\u6c0f\u9670\u6027\u83cc\uff0c\u690d\u7269\u5728\u7f3a\u6c2e\u6642\uff0c\u6839\u90e8\u6703\u5206\u6ccc\u751c\u83dc\u9e7c(betaine)\u3001\u985e\u9ec3\u916e(flavonoid)\u8207\u7570\u9ec3\u916e(isoflavonoid)\u7b49\u7269\u8cea\u4f86\u5438\u5f15\u6839\u7624\u83cc\uff1b\u800c\u6839\u7624\u83cc\u53d7\u5230\u5438\u5f15\u5f8c\uff0c\u6703\u5206\u6ccc\u7531\u7cd6\u8207\u8102\u80aa\u9178\u69cb\u6210\u7684Nod factor(\u5982\u4e0b\u5716)\uff0c\u4f5c\u7528\u5728\u690d\u7269\u7684\u6839\u6bdb\u4e0a\uff0c\u4f7f\u81ea\u5df1\u66f4\u5bb9\u6613\u9644\u8457\u3002<\/p>\n<figure id=\"attachment_20669\" aria-describedby=\"caption-attachment-20669\" style=\"width: 400px\" class=\"wp-caption aligncenter\"><img loading=\"lazy\" decoding=\"async\" class=\"wp-image-20669\" src=\"http:\/\/case.ntu.edu.tw\/blog\/wp-content\/uploads\/2015\/02\/NodSm-IV_AcC16-2S.svg_-1024x837.png\" alt=\"\u5716\u7247\u4f86\u6e90\uff1awiki\" width=\"400\" height=\"327\" srcset=\"https:\/\/case.ntu.edu.tw\/blog\/wp-content\/uploads\/2015\/02\/NodSm-IV_AcC16-2S.svg_-1024x837.png 1024w, https:\/\/case.ntu.edu.tw\/blog\/wp-content\/uploads\/2015\/02\/NodSm-IV_AcC16-2S.svg_-300x245.png 300w, https:\/\/case.ntu.edu.tw\/blog\/wp-content\/uploads\/2015\/02\/NodSm-IV_AcC16-2S.svg_-600x491.png 600w, https:\/\/case.ntu.edu.tw\/blog\/wp-content\/uploads\/2015\/02\/NodSm-IV_AcC16-2S.svg_-800x654.png 800w, https:\/\/case.ntu.edu.tw\/blog\/wp-content\/uploads\/2015\/02\/NodSm-IV_AcC16-2S.svg_.png 1280w\" sizes=\"(max-width: 400px) 100vw, 400px\" \/><figcaption id=\"caption-attachment-20669\" class=\"wp-caption-text\">\u5716\u7247\u4f86\u6e90\uff1awiki<\/figcaption><\/figure>\n<p>\u63a5\u8457\u5c31\u958b\u59cb\u6839\u7624\u7684\u5f62\u6210\u3002\u76ee\u524d\u77e5\u9053\uff0c\u7a76\u7adf\u4e00\u68f5\u690d\u7269\u80fd\u5920\u5f62\u6210\u591a\u5c11\u6839\u7624\uff0c\u5176\u5be6\u662f\u7d93\u904e\u7cbe\u5bc6\u7684\u8abf\u7bc0\u6a5f\u5236\u7684\uff0c\u53ef\u80fd\u4e5f\u8207\u7d30\u80de\u5206\u88c2\u7d20(cytokinin)\u6709\u95dc(\u53c33)\u3002<\/p>\n<p>\u81f3\u65bc\u70ba\u4ec0\u9ebc\u8981\u5f62\u6210\u6839\u7624\u5462\uff1f\u5176\u5be6\u662f\u56e0\u70ba\u5927\u6c23\u4e2d\u7684\u6c2e\u662f\u975e\u5e38\u975e\u5e38\u7a69\u5b9a\u7684\u4e09\u9375\u7d50\u69cb\uff0c\u8981\u6253\u7834\u4e00\u5206\u5b50\u7684\u6c2e\u9700\u898116\u500bATP\uff1b\u5728\u9019\u9ebc\u9ad8\u7684\u80fd\u91cf\u9700\u6c42\u4e0b\uff0c\u6839\u7624\u83cc\u4e2d\u7684\u56fa\u6c2e\u9175\u7d20(nitrogenase)\u7121\u6cd5\u5728\u6c27\u6c23\u5b58\u5728\u4e0b\u9032\u884c\u56fa\u6c2e\u53cd\u61c9\u3002\u4e8b\u5be6\u4e0a\uff0c\u56fa\u6c2e\u9175\u7d20\u53ea\u8981\u63a5\u89f8\u5230\u6c27\u6c23\u5927\u7d04\u56db\u5206\u534a\u9418\uff0c\u6d3b\u6027\u5c31\u5269\u4e0b\u4e0d\u5230\u767e\u5206\u4e4b\u4e00\u4e86(\u53c34)\u3002<\/p>\n<p>\u6839\u7624\uff0c\u662f\u690d\u7269\u70ba\u4e86\u8b93\u6839\u7624\u83cc\u53ef\u4ee5\u56fa\u6c2e\u6240\u5f62\u6210\u7684\u7279\u6b8a\u69cb\u9020(\u5982\u4e0b\u5716)\u3002\u6839\u7624\u7684\u8868\u9762\u6709\u539a\u539a\u7684\u8868\u76ae\uff08\u4e0b\u5716\u5de6\u4e0b\u89d2\uff09\u9694\u7d55\u7a7a\u6c23\uff0c\u4f7f\u6839\u7624\u5167\u90e8\u5448\u73fe\u5fae\u6c27(microaerobic)\u7684\u72c0\u614b\u3002\u5167\u90e8\u7684\u6839\u7624\u83cc\u5728\u5fae\u6c27\u72c0\u614b\u4e0b\u5448\u73fe\u7279\u6b8a\u7684\u578b\u614b\uff0c\u88ab\u7a31\u70ba\u300c\u985e\u7d30\u83cc\u300d(bacteroid)\uff1b\u800c\u6839\u7624\u5167\u90e8\u7684\u690d\u7269\u7d30\u80de\u4ee5\u53ca\u6839\u7624\u83cc\u6240\u9700\u8981\u7684\u6c27\u6c23\uff0c\u5247\u662f\u7531\u300c\u5171\u751f\u8840\u7d05\u7d20\u300d(symbiotic\uff0csHbs)\u4f86\u4f9b\u61c9\u3002<\/p>\n<figure id=\"attachment_20670\" aria-describedby=\"caption-attachment-20670\" style=\"width: 400px\" class=\"wp-caption aligncenter\"><img loading=\"lazy\" decoding=\"async\" class=\"wp-image-20670\" src=\"http:\/\/case.ntu.edu.tw\/blog\/wp-content\/uploads\/2015\/02\/1280px-Root-nodule01-1024x756.jpg\" alt=\"\u6839\u7624\u7684\u5207\u9762\u3002\u5716\u7247\u4f86\u6e90\uff1awiki\" width=\"400\" height=\"295\" srcset=\"https:\/\/case.ntu.edu.tw\/blog\/wp-content\/uploads\/2015\/02\/1280px-Root-nodule01-1024x756.jpg 1024w, https:\/\/case.ntu.edu.tw\/blog\/wp-content\/uploads\/2015\/02\/1280px-Root-nodule01-300x221.jpg 300w, https:\/\/case.ntu.edu.tw\/blog\/wp-content\/uploads\/2015\/02\/1280px-Root-nodule01-600x443.jpg 600w, https:\/\/case.ntu.edu.tw\/blog\/wp-content\/uploads\/2015\/02\/1280px-Root-nodule01-800x591.jpg 800w, https:\/\/case.ntu.edu.tw\/blog\/wp-content\/uploads\/2015\/02\/1280px-Root-nodule01.jpg 1280w\" sizes=\"(max-width: 400px) 100vw, 400px\" \/><figcaption id=\"caption-attachment-20670\" class=\"wp-caption-text\">\u6839\u7624\u7684\u5207\u9762\u3002\u5716\u7247\u4f86\u6e90\uff1awiki<\/figcaption><\/figure>\n<p>\u5176\u5be6\uff0c\u9664\u4e86\u6839\u7624\u83cc\u5c6c\u4ee5\u5916\uff0c\u80fd\u5920\u8207\u690d\u7269\u5f62\u6210\u5171\u751f\u95dc\u4fc2\u4f86\u56fa\u6c2e\u7684\uff0c\u9084\u5305\u62ec\u4e86\u00a0<em>Bradyrhizobium(<\/em>\u6162\u751f\u6839\u7624\u83cc\u5c6c),\u00a0<em>Sinorhizobium(<\/em>\u4e2d\u83ef\u6839\u7624\u83cc\u5c6c),\u00a0<em>Azorhizobium<\/em>,\u00a0<em>Mesorhizobium(<\/em>\u4e2d\u6162\u751f\u6839\u7624\u83cc\u5c6c),\u00a0<em>Burkholderia<\/em><em>\u00a0(<\/em>\u985e\u9f3b\u75bd\u83cc\u5c6c)\u8207\u00a0<em>Cupriavidus(<\/em>\u8caa\u9285\u83cc\u5c6c)\u7b49\uff0c\u56ca\u62ec\u4e86alpha- \u8207 beta- proteobacteria(\u03b1\u8207\u03b2\u8b8a\u5f62\u83cc)\u3002\u4e0d\u904e\uff0c\u7531\u65bc\u9019\u4e9b\u83cc\u7684<em>nod<\/em>\u57fa\u56e0\u76f8\u4f3c\u6027\u6975\u9ad8\uff0c\u63a8\u60f3\u61c9\u8a72\u662f\u900f\u904e\u8cea\u9ad4\u5e73\u884c\u8f49\u79fb(horizontal transfer)\u7684\u7d50\u679c\u3002<\/p>\n<p>\u76f4\u52301909\u5e74\u54c8\u4f2f(Fritz Haber)\u767c\u660e\u5de5\u696d\u56fa\u6c2e\u4e4b\u524d\uff0c\u8fb2\u696d\u7684\u6c2e\u80a5\u4f86\u6e90\uff0c\u9664\u4e86\u5806\u80a5\u4ee5\u5916\uff0c\u5c31\u662f\u7531\u667a\u5229\u751f\u7522\u7684\u751f\u785d(Caliche\uff0c\u6210\u5206\u70ba\u785d\u9178\u9209)\u3002\u5728\u5de5\u696d\u56fa\u6c2e\u767c\u660e\u4e4b\u5f8c\uff0c\u667a\u5229\u7684\u751f\u785d\u5c31\u6c92\u4eba\u8981\u4e86\uff1a\u7522\u91cf\u75311925\u5e74\u7684\u6bcf\u567845\u7f8e\u91d1\u843d\u52301934\u5e74\u7684\u6bcf\u567819\u7f8e\u91d1\uff0c\u800c\u4e14\u7531250\u842c\u5678\u964d\u523080\u842c\u5678\u3002\u79d1\u6280\u7684\u9032\u6b65\uff0c\u96d6\u7136\u9020\u798f\u4e86\u8a31\u591a\u4eba\uff0c\u4f46\u662f\u5c0d\u7576\u5e74\u5931\u696d\u7684\u56db\u842c\u4e94\u5343\u516b\u767e\u591a\u540d\u54e1\u5de5\u4ee5\u53ca\u4ed6\u5011\u7684\u5bb6\u5ead\u4f86\u8aaa(\u53c35)\uff0c\u9084\u771f\u7684\u5f88\u96e3\u8aaa\u662f\u798f\u5462\uff01<\/p>\n<p>\u9664\u4e86\u8c46\u79d1\u690d\u7269\u4ee5\u5916\uff0c\u80fd\u8207\u56fa\u6c2e\u5fae\u751f\u7269\u5171\u751f\u7684\u690d\u7269\u5305\u62ec\u5c0f\u982d\u6728\u9ebb\u9ec3(<em>Casuarina oligodon<\/em>\uff0c\u8207Frankia\u5c6c\u83cc\u5171\u751f)\u3001\u6eff\u6c5f\u7d05(<em>Azolla<\/em>\uff0c\u8207<em>Anabaena<\/em>\u5c6c\u7684\u85cd\u7da0\u83cc\u5171\u751f)\uff1b\u9019\u5169\u7a2e\u690d\u7269\u90fd\u5728\u8fb2\u696d\u4e0a\u88ab\u5ee3\u6cdb\u7684\u61c9\u7528\uff1a\u5c0f\u982d\u6728\u9ebb\u9ec3\u5728\u65b0\u5e7e\u5167\u4e9e\u3001\u6eff\u6c5f\u7d05\u5247\u662f\u5728\u4e2d\u570b\u5357\u65b9\u4ee5\u53ca\u53f0\u7063\u7684\u6c34\u7a3b\u7530\u4e2d\u505a\u70ba\u5929\u7136\u6c2e\u80a5\u7684\u4f86\u6e90\u3002\u53ea\u53ef\u60dc\u96a8\u8457\u54c8\u4f2f\u6cd5\u7684\u767c\u660e\u800c\u4f86\u7684\u5316\u80a5\uff0c\u8981\u5728\u6c34\u7a3b\u7530\u4e2d\u770b\u5230\u6eff\u6c5f\u7d05\u5df2\u7d93\u975e\u5e38\u4e0d\u5bb9\u6613\u4e86\uff01<\/p>\n<figure id=\"attachment_20677\" aria-describedby=\"caption-attachment-20677\" style=\"width: 558px\" class=\"wp-caption aligncenter\"><img loading=\"lazy\" decoding=\"async\" class=\"size-full wp-image-20677\" src=\"http:\/\/case.ntu.edu.tw\/blog\/wp-content\/uploads\/2015\/03\/524230_3224801061338_1306836142_n.jpg\" alt=\"\u6709\u6a5f\u6c34\u7a3b\u7530\u88e1\u7684\u6eff\u6c5f\u7d05\uff08\u4f5c\u8005\u651d\u65bc\u6148\u6fdf\u57fa\u91d1\u6703\u5927\u611b\u8fb2\u5834\uff09\" width=\"558\" height=\"417\" srcset=\"https:\/\/case.ntu.edu.tw\/blog\/wp-content\/uploads\/2015\/03\/524230_3224801061338_1306836142_n.jpg 558w, https:\/\/case.ntu.edu.tw\/blog\/wp-content\/uploads\/2015\/03\/524230_3224801061338_1306836142_n-300x224.jpg 300w\" sizes=\"(max-width: 558px) 100vw, 558px\" \/><figcaption id=\"caption-attachment-20677\" class=\"wp-caption-text\">\u6709\u6a5f\u6c34\u7a3b\u7530\u88e1\u7684\u6eff\u6c5f\u7d05\uff08\u4f5c\u8005\u651d\u65bc\u6148\u6fdf\u57fa\u91d1\u6703\u5927\u611b\u8fb2\u5834\uff09<\/figcaption><\/figure>\n<p><strong>\u53c3\u8003\u6587\u737b\uff1a<\/strong><br \/>\n1.2014\/8\/17.\u00a0<a href=\"http:\/\/susan-plant-kingdom.blogspot.tw\/2014\/08\/blog-post_11.html\" target=\"_blank\" rel=\"noopener noreferrer\">\u539f\u4f86\u79e6\u671d\u3001\u6f22\u671d\u7684\u4e3b\u98df\u662f\u5c0f\u7c73\uff01<\/a>\u8001\u8449\u7684\u690d\u7269\u738b\u570b\u3002<br \/>\n2.Ann M. Hirsch 2009.\u00a0<a href=\"https:\/\/www.mcdb.ucla.edu\/Research\/Hirsch\/imagesb\/HistoryDiscoveryN2fixingOrganisms.pdf\" target=\"_blank\" rel=\"noopener noreferrer\">Brief History of the Discovery of Nitrogen-fixing Organisms.<\/a><br \/>\n3.2014\/9\/21.\u00a0<a href=\"http:\/\/susan-plant-kingdom.blogspot.tw\/2014\/09\/root-nodule.html\" target=\"_blank\" rel=\"noopener noreferrer\">\u8c46\u79d1\u690d\u7269\u5982\u4f55\u4fdd\u6301\u4e0d\u591a\u4e0d\u5c11\u7684\u6839\u7624\uff08root nodule\uff09<\/a>\uff1f\u8001\u8449\u7684\u690d\u7269\u738b\u570b\u3002<br \/>\n4.Taiz and Zeiger. Plant Physiology. 5th ed.<br \/>\n5.Wikipedia.\u00a0<a href=\"http:\/\/en.wikipedia.org\/wiki\/Fritz_Haber\uff3f\" target=\"_blank\" rel=\"noopener noreferrer\">Fritz Haber<\/a>.<\/p>\n<p><span style=\"color: #333399;\"><strong>--<\/strong><\/span><br \/>\n<span style=\"color: #333399;\"><strong> \u4f5c\u8005\uff1a\u8449\u7da0\u8212\u3000\u6148\u6fdf\u5927\u5b78\u751f\u547d\u79d1\u5b78\u7cfb\u52a9\u7406\u6559\u6388\uff0c\u79d1\u6559\u4e2d\u5fc3\u7279\u7d04\u5beb\u624b\uff0c\u5f9e\u4e8b\u79d1\u666e\u6587\u7ae0\u5beb\u4f5c\u3002<\/strong><\/span><\/p>\n<p>&nbsp;<\/p>\n<p><a href=\"https:\/\/line.me\/R\/ti\/p\/%40fcx6182w\" target=\"_blank\" rel=\"noopener noreferrer\"><img loading=\"lazy\" decoding=\"async\" class=\"alignnone\" src=\"https:\/\/scdn.line-apps.com\/n\/line_add_friends\/btn\/zh-Hant.png\" alt=\"\u52a0\u5165\u597d\u53cb\" width=\"212\" height=\"36\" border=\"0\" \/><\/a><\/p>\n","protected":false},"excerpt":{"rendered":"<p>\u64b0\u6587\uff5c\u8449\u7da0\u8212 \u81ea\u5f9e\u4eba\u985e\u5728\u4e00\u842c\u4e00\u5343\u5e74\u524d\u5b78\u6703<\/p>\n","protected":false},"author":20,"featured_media":20664,"comment_status":"closed","ping_status":"closed","sticky":false,"template":"","format":"standard","meta":{"footnotes":""},"categories":[3293,3100],"tags":[1642,1660,1466,1543,1170],"aioseo_notices":[],"jetpack_featured_media_url":"https:\/\/case.ntu.edu.tw\/blog\/wp-content\/uploads\/2015\/02\/Soybean.USDA_.jpg","_links":{"self":[{"href":"https:\/\/case.ntu.edu.tw\/blog\/index.php?rest_route=\/wp\/v2\/posts\/20895"}],"collection":[{"href":"https:\/\/case.ntu.edu.tw\/blog\/index.php?rest_route=\/wp\/v2\/posts"}],"about":[{"href":"https:\/\/case.ntu.edu.tw\/blog\/index.php?rest_route=\/wp\/v2\/types\/post"}],"author":[{"embeddable":true,"href":"https:\/\/case.ntu.edu.tw\/blog\/index.php?rest_route=\/wp\/v2\/users\/20"}],"replies":[{"embeddable":true,"href":"https:\/\/case.ntu.edu.tw\/blog\/index.php?rest_route=%2Fwp%2Fv2%2Fcomments&post=20895"}],"version-history":[{"count":3,"href":"https:\/\/case.ntu.edu.tw\/blog\/index.php?rest_route=\/wp\/v2\/posts\/20895\/revisions"}],"predecessor-version":[{"id":31517,"href":"https:\/\/case.ntu.edu.tw\/blog\/index.php?rest_route=\/wp\/v2\/posts\/20895\/revisions\/31517"}],"wp:featuredmedia":[{"embeddable":true,"href":"https:\/\/case.ntu.edu.tw\/blog\/index.php?rest_route=\/wp\/v2\/media\/20664"}],"wp:attachment":[{"href":"https:\/\/case.ntu.edu.tw\/blog\/index.php?rest_route=%2Fwp%2Fv2%2Fmedia&parent=20895"}],"wp:term":[{"taxonomy":"category","embeddable":true,"href":"https:\/\/case.ntu.edu.tw\/blog\/index.php?rest_route=%2Fwp%2Fv2%2Fcategories&post=20895"},{"taxonomy":"post_tag","embeddable":true,"href":"https:\/\/case.ntu.edu.tw\/blog\/index.php?rest_route=%2Fwp%2Fv2%2Ftags&post=20895"}],"curies":[{"name":"wp","href":"https:\/\/api.w.org\/{rel}","templated":true}]}}