{"id":19308,"date":"2014-10-17T00:58:44","date_gmt":"2014-10-16T16:58:44","guid":{"rendered":"https:\/\/case.ntu.edu.tw\/blog\/?p=19308"},"modified":"2017-07-25T10:57:27","modified_gmt":"2017-07-25T02:57:27","slug":"%e3%80%90%e7%94%9f%e7%89%a9%e5%a5%a7%e5%a6%99%e3%80%91%e4%ba%ba%e7%ae%97%e4%b8%8d%e5%a6%82%e5%a4%a9%e7%ae%97%e4%b9%8b%e5%85%89%e5%90%88%e4%bd%9c%e7%94%a8%e6%94%b9%e9%80%b2","status":"publish","type":"post","link":"https:\/\/case.ntu.edu.tw\/blog\/?p=19308","title":{"rendered":"\u3010\u751f\u7269\u5967\u5999\u3011\u4eba\u7b97\u4e0d\u5982\u5929\u7b97\u4e4b\u5149\u5408\u4f5c\u7528\u6539\u9032"},"content":{"rendered":"

\u25a0\u5149\u5408\u4f5c\u7528\uff08photosynthesis\uff09\u88e1\u9762\u6700\u91cd\u8981\u7684\u9175\u7d20\u5c31\u662f\u7c21\u7a31Rubisco\u7684\u9175\u7d20\u4e86\u3002<\/strong><\/span><\/p>\n

\"Rubisco\u3002\u5716\u7247\u4f86\u6e90\uff1awiki\"
Rubisco\u3002\u5716\u7247\u4f86\u6e90\uff1awiki<\/figcaption><\/figure>\n

\u64b0\u6587\uff5c<\/strong>\u8449\u7da0\u8212<\/p>\n

\u9019\u500b\u9175\u7d20\u6709\u500b\u5f88\u9577\u7684\u540d\u5b57\uff0cD-ribulose-1,5-bisphosphate carboxylase\/oxygenase\uff0c\u56e0\u70ba\u9175\u7d20\u547d\u540d\u7684\u901a\u5247\u662f\u300c\u53d7\u8cea\u6216\u7522\u7269\u540d\u7a31\u300d\uff08D-ribulose-1,5-bisphosphate\uff09\uff0b\u300c\u53cd\u61c9\u985e\u578b\u300d\uff08carboxylase\/oxygenase\uff09\uff0c\u6240\u4ee5\u5c31\u6709\u500b\u300c\u843d\u843d\u9577\u300d\u7684\u540d\u5b57\u3002<\/p>\n

\u5f9e\u5b83\u7684\u540d\u5b57\u53ef\u4ee5\u770b\u5230\uff0c\u9019\u500b\u9175\u7d20\u6709\u5169\u7a2e\u53cd\u61c9\u985e\u578b\uff1a\u4e00\u500b\u662f\u6211\u5011\u719f\u6089\u7684\u53cd\u61c9\u985e\u578b\uff0c\u5c07\u4e8c\u6c27\u5316\u78b3\u8207D-ribulose-1,5-bisphosphate\uff08RuBP\uff09\u7d50\u5408\uff0c\u7522\u751f\u5169\u500b\u5206\u5b50\u7684\u5e36\u6709\u78f7\u9178\u6839\u7684\u4e09\u78b3\u5316\u5408\u72693-phosphoglycreate \uff083-PGA\uff09\uff0c\u9019\u500b\u53cd\u61c9\u662f\u6240\u8b02\u7684\u7fa7\u57fa\u5316\uff08carboxylation\uff09\uff1b\u53e6\u4e00\u500b\u5247\u662f\u5c07RuBP\u6c27\u5316\uff0c\u7522\u751f\u4e00\u5206\u5b50\u76843-PGA\u4ee5\u53ca\u4e00\u5206\u5b50\u76842 - \u78f7\u9178\u4e59\u9187\u9178\uff082-phosphoglycolate\uff09\u30022 - \u78f7\u9178\u4e59\u9187\u9178\u5c0d\u690d\u7269\u7121\u7528\uff0c\u5fc5\u9700\u6d88\u8017\u80fd\u91cf\u4f86\u56de\u6536\u5b83\uff0c\u56de\u6536\u7684\u904e\u7a0b\u88ab\u7a31\u70ba\u5149\u547c\u5438\uff08photorespiration\uff09\u4f5c\u7528\u3002<\/p>\n

\u56e0\u70ba\u5149\u547c\u5438\u4f5c\u7528\u6703\u6d88\u8017\u80fd\u91cf\uff0c\u5c0d\u690d\u7269\u4f86\u8aaa\u7576\u7136\u662f\u6709\u5bb3\u7121\u76ca\uff1b\u4f46\u662f\u504f\u504fRubisco\u5c31\u662f\u6c92\u8fa6\u6cd5\u628a\u5c0d\u6c27\u6c23\u7684\u89aa\u548c\u529b\u5b8c\u5168\u53bb\u9664\u3002\u7531\u65bcRubisco\u53ea\u80fd\u5229\u7528\u6eb6\u65bc\u6c34\u7684\u6c23\u9ad4\uff08\u4e0d\u8ad6\u662f\u6c27\u6216\u4e8c\u6c27\u5316\u78b3\uff09\uff0c\u7576\u6eab\u5ea6\u4e0a\u6607\u6642\u6c23\u9ad4\u5c0d\u6c34\u7684\u6eb6\u89e3\u5ea6\u6703\u4e0b\u964d\uff1b\u800c\u4e8c\u6c27\u5316\u78b3\u56e0\u70ba\u5206\u5b50\u91cf\u6bd4\u6c27\u5927\uff0c\u6240\u4ee5\u6eb6\u89e3\u5ea6\u4e0b\u964d\u5f97\u66f4\u660e\u986f\uff0c\u9020\u6210\u5728\u6c23\u6eab\u4e0a\u6607\u7684\u6642\u5019\uff0c\u5149\u547c\u5438\u4f5c\u7528\u6703\u8b8a\u5f97\u66f4\u5287\u70c8\u3002<\/p>\n

\u5c0d\u65bc\u6eab\u5e36\u690d\u7269\u6216\u8a31\u9019\u500b\u554f\u984c\u53ea\u662f\u5c0f\u4e8b\u60c5\uff0c\u4f46\u662f\u5c0d\u65bc\u71b1\u5e36\u8207\u4e9e\u71b1\u5e36\u7684\u690d\u7269\u4f86\u8aaa\uff0c\u5149\u547c\u5438\u4f5c\u7528\u662f\u4ed6\u5011\u4e00\u5e74\u5230\u982d\u90fd\u8981\u8655\u7406\u7684\u554f\u984c\u3002<\/p>\n

\u65bc\u662f\uff0c\u67d0\u4e9b\u71b1\u5e36\u8207\u4e9e\u71b1\u5e36\u7684\u690d\u7269\uff08\u5982\uff1a\u7389\u7c73\uff09\uff0c\u6f14\u5316\u51fa\u4e86C4\u4ee3\u8b1d\u3002C4\u4ee3\u8b1d\u4e26\u6c92\u6709\u628aRubisco\u7d66\u6d88\u6ec5\uff0c\u800c\u662f\u5728Rubisco\u7684\u53cd\u61c9\u524d\u9762\uff0c\u52a0\u5165\u4e86\u4e00\u500b\u6fc3\u7e2e\u4e8c\u6c27\u5316\u78b3\u7684\u6a5f\u5236\uff08CO2-concentrating mechanism\uff0cCCM\uff09\u3002\u540c\u6642\uff0c\u57f7\u884cC4\u4ee3\u8b1d\u7684\u690d\u7269\u628aCCM\u8207\u5305\u542bRubisco\u5728\u5167\u7684\u6574\u500b\u5361\u723e\u6587\u5faa\u74b0\uff08Calvin cycle\uff09\u5206\u958b\uff0c\u5c07\u5361\u723e\u6587\u5faa\u74b0\u95dc\u5230\u9ad3\u9798\u7d30\u80de\uff08bundle-sheath cell\uff09\u4e2d\uff0c\u6e1b\u5c11Rubisco\u8207\u6c27\u6c23\u7684\u63a5\u89f8\uff0c\u4f7f\u5f97C4\u4ee3\u8b1d\u53ef\u4ee5\u6210\u529f\u5730\u5c07\u5149\u547c\u5438\u4f5c\u7528\u7d66\u6d88\u6ec5\u3002<\/p>\n

\"C4\u690d\u7269\u7684\u8449\u7247\u6a6b\u5207\u9762\u3002
C4\u690d\u7269\u7684\u8449\u7247\u6a6b\u5207\u9762\u3002
\u7da0\u8272\u90e8\u5206\u662f\u8449\u8089\u7d30\u80de\uff08mesophyll\uff09\uff0c\u570d\u7e5e\u8457\u4e2d\u5fc3\u7684
\u7dad\u7ba1\u675f\uff08\u7d05\u8272\uff09\u7684\u7d2b\u8272\u90e8\u5206\u5c31\u662f\u9ad3\u9798\u7d30\u80de\u3002
\u5716\u7247\u4f86\u6e90\uff1awiki<\/figcaption><\/figure>\n

\u56e0\u70baC4\u4ee3\u8b1d\u6210\u529f\u5730\u6d88\u6ec5\u4e86\u5149\u547c\u5438\u4f5c\u7528\uff0c\u65bc\u662f\u4ed6\u5011\u53ef\u4ee5\u5728\u590f\u5929\u6642\u7121\u8996\u65bc\u5916\u754c\u7684\u9ad8\u6eab\uff0c\u4ecd\u7136\u7e7c\u7e8c\u5feb\u6a02\u7684\u751f\u9577\uff1b\u4f46\u662fC4\u690d\u7269\u7562\u7adf\u662f\u5c11\u6578\uff0c\u4e16\u754c\u4e0a\u4e3b\u8981\u7684\u7ce7\u98df\u4f5c\u7269\uff0c\u9664\u4e86\u7389\u7c73\u662fC4\u690d\u7269\uff0c\u5176\u4ed6\u7684\u5c0f\u9ea5\u3001\u5927\u9ea5\u3001\u7a3b\u7c73\u3001\u5927\u8c46...\u5168\u90fd\u662f\u4e0d\u5177\u6709CCM\u7684C3\u690d\u7269\u3002<\/p>\n

\u8fd1\u5e74\u4f86\uff0c\u56e0\u70ba\u6c23\u5019\u8b8a\u9077\u4ee5\u53ca\u4eba\u53e3\u589e\u9577\uff0c\u4f7f\u5f97\u79d1\u5b78\u5bb6\u5011\u60f3\u5230\uff0c\u5982\u679c\u53ef\u4ee5\u6539\u9032\u9019\u4e9bC3\u690d\u7269\u7684\u5149\u5408\u4f5c\u7528\uff0c\u8b93\u4ed6\u5011\u4e0d\u8981\u6d88\u8017\u9019\u9ebc\u591a\u80fd\u91cf\u5728\u5149\u547c\u5438\u4f5c\u7528\u4e0a\uff0c\u61c9\u8a72\u53ef\u4ee5\u5927\u5927\u5730\u63d0\u5347\u7522\u91cf\u5427\uff1f\u53ef\u60dc\u7684\u662f\uff0c\u8981\u628aC3\u690d\u7269\u6539\u6210C4\u690d\u7269\u4e26\u975e\u5bb9\u6613\u7684\u4e8b\uff0c\u7562\u7adf\u9019\u727d\u6d89\u5230\u89e3\u5256\u5b78\u4e0a\u7684\u69cb\u9020\u6539\u8b8a\uff1b\u65bc\u662f\u6709\u4e9b\u79d1\u5b78\u5bb6\u60f3\u5230\uff0c\u662f\u5426\u80fd\u5920\u53ea\u628a\u8ddfCCM\u76f8\u95dc\u7684\u9175\u7d20\u9001\u9032\u8449\u7da0\u9ad4\u4e2d\uff0c\u540c\u6642\u4e5f\u53d6\u4ee3\u539f\u6709\u7684Rubisco\uff0c\u770b\u770b\u662f\u5426\u80fd\u4f7f\u5f97C3\u690d\u7269\u6709\u4e9b\u6539\u8b8a\u3002<\/p>\n

\u6700\u8fd1\u5728\u5eb7\u4e43\u723e\u5927\u5b78\uff08Cornell University\uff09\u7684\u7814\u7a76\u5718\u968a\u767c\u8868\u5728\u300c\u81ea\u7136\u300d\u671f\u520a\u4e0a\u7684\u8ad6\u6587\uff0c\u5c31\u662f\u9019\u500b\u60f3\u6cd5\u7684\u9ad4\u73fe\u3002\u7814\u7a76\u5718\u968a\u5c07\u85cd\u7da0\u85fb\uff08cyanobacteria\uff09\u7684Rubisco\u4ee5\u53caCCM\u76f8\u95dc\u9175\u7d20\uff0c\u6574\u5806\u9001\u9032\u83f8\u8349\uff08C3\u690d\u7269\uff09\u7684\u8449\u7da0\u9ad4\u88e1\u9762\uff0c\u60f3\u8981\u770b\u770b\u662f\u4e0d\u662f\u80fd\u6539\u9032\u83f8\u8349\u7684\u5149\u5408\u4f5c\u7528\u6548\u7387\u3002<\/p>\n

\u7b49\u4e00\u4e0b\uff01\u600e\u4e0d\u662f\u653e\u7389\u7c73\u6216\u5176\u4ed6\u9ad8\u7b49C4\u690d\u7269\uff1f\u53cd\u800c\u653e\u4e86\u85cd\u7da0\u85fb\uff1f<\/p>\n

\u7b46\u8005\u60f3\uff0c\u5982\u679c\u80fd\u653e\u9ad8\u7b49C4\u690d\u7269\u7684\u9175\u7d20\u7576\u7136\u662f\u4e0a\u4e0a\u5927\u5409\uff0c\u4e0d\u904e\u727d\u6d89\u5230\u7684\u57fa\u56e0\u592a\u591a\u4e86\uff0c\u8981\u653e\u9032\u53bb\u53ef\u80fd\u4e0d\u5bb9\u6613\u3002<\/p>\n

\"C4\u4ee3\u8b1d\u4e4b\u4e00\u7a2e\u3002\u5716\u7247\u4f86\u6e90\uff1awiki<\/a>
C4\u4ee3\u8b1d\u4e4b\u4e00\u7a2e\u3002\u5716\u7247\u4f86\u6e90\uff1awiki<\/figcaption><\/figure>\n

\u5927\u5bb6\u53ef\u4ee5\u53c3\u8003\u4e0a\u9762\u7684\u5716\uff0c\u5de6\u908a\u9ec3\u8272\u7d30\u80de\u88e1\u9762\u6240\u6709\u7684\u53cd\u61c9\uff0c\u5c31\u662f\u9ad8\u7b49C4\u690d\u7269\u7684CCM\u3002\u8449\u7da0\u9ad4\u8f49\u6b96\u662f\u6bd4\u690d\u7269\u7d30\u80de\u8f49\u6b96\u96e3\u5341\u500d\u4ee5\u4e0a\u7684\u6280\u8853\uff0c\u7136\u5f8c\u9084\u8981\u653e\u9019\u9ebc\u591a\u500b\u57fa\u56e0...\u7b46\u8005\u60f3\uff0c\u9019\u662f\u5eb7\u4e43\u723e\u7814\u7a76\u5718\u968a\u6700\u5f8c\u9078\u4e86\u85cd\u7da0\u85fb\u7684\u539f\u56e0\u3002\u96d6\u7136\u4ed6\u4e0d\u662fC4\u690d\u7269\uff0c\u4f46\u662f\u4ed6\u4e5f\u6709CCM\uff0c\u56e0\u6b64\u4e5f\u5177\u5099CCM\u7684\u9175\u7d20\uff1b\u800c\u7531\u65bc\u4ed6\u662f\u8f03\u4f4e\u7b49\u7684\u690d\u7269\uff0c\u6240\u4ee5CCM\u7684\u9175\u7d20\u6210\u54e1\u6c92\u6709\u90a3\u9ebc\u591a...<\/p>\n

\u65bc\u662f\uff0c\u7814\u7a76\u5718\u968a\u5011\u5c07\u85cd\u7da0\u85fb\u7684Rubisco\u4ee5\u53caCCM\u9175\u7d20\u8f49\u6b96\u5230\u83f8\u8349\u7684\u8449\u7da0\u9ad4\u88e1\u9762\u53bb\uff0c\u540c\u6642\u4e5f\u5c07\u83f8\u8349\u672c\u4f86\u7684Rubisco\u780d\u6389\u3002<\/p>\n

\u807d\u8d77\u4f86\u597d\u50cf\u90fd\u5f88\u4e0d\u932f\uff0c\u537b\u6c92\u60f3\u5230\u85cd\u7da0\u85fb\u7684Rubisco\u5c0d\u4e8c\u6c27\u5316\u78b3\u7684\u89aa\u548c\u529b\uff0c\u6bd4\u83f8\u8349\u7684Rubisco\u9084\u8981\u4f4e\uff01\u56e0\u70ba\u9019\u6a23\uff0c\u4f7f\u5f97\u8f49\u6b96\u7684\u83f8\u8349\u7121\u6cd5\u5728400ppm\u7684\u4e8c\u6c27\u5316\u78b3\u7684\u74b0\u5883\u4e0b\u751f\u9577\uff08\u9019\u662f\u76ee\u524d\u6b63\u5e38\u5927\u6c23\u7684\u4e8c\u6c27\u5316\u78b3\u6fc3\u5ea6\uff09\uff0c\u5fc5\u9700\u8981\u63d0\u9ad8\u4e8c\u5341\u5e7e\u500d\uff0c\u52309000ppm\u624d\u9577\u5f97\u8d77\u4f86\uff0c\u800c\u4e14\u9084\u9577\u5f97\u5f88\u6162\u5462...<\/p>\n

\u600e\u9ebc\u6703\u9019\u6a23\u5462\uff1f\u539f\u4f86\uff0c\u96d6\u7136\u9019\u4e9b\u5177\u6709CCM\u7684\u690d\u7269\uff0c\u5305\u62ecC4\u690d\u7269\u3001\u85fb\u985e\u3001\u4ee5\u53ca\u85cd\u7da0\u85fb\uff0c\u96d6\u7136\u4ed6\u5011\u7684Rubisco\u53cd\u61c9\u901f\u7387\u6bd4C3\u690d\u7269\u7684\u8981\u5feb\u5f97\u591a\uff0c\u4f46\u662f\u537b\u7121\u6cd5\u5728\u52a0\u5feb\u901f\u7387\u8207\u9ad8\u89aa\u548c\u529b\uff08\u4e5f\u5c31\u662f\u4f4eKm\u503c\uff09\u4e4b\u9593\u517c\u5f97\u3002\u65bc\u662f\u9020\u6210\u8f49\u6b96\u7684\u83f8\u8349\u7121\u6cd5\u5728\u5927\u6c23\u74b0\u5883\u7684\u4e8c\u6c27\u5316\u78b3\u4e2d\u751f\u9577\uff0c\u56e0\u70ba\u5b83\u6293\u4e0d\u5230\u4e8c\u6c27\u5316\u78b3...<\/p>\n

\u9019\u771f\u7684\u662f\u8b93\u4eba\u89ba\u5f97\u907a\u61be\u842c\u5206\uff0c\u4e0d\u904e\uff0c\u8981\u9b5a\u8207\u718a\u638c\u517c\u5f97\uff0c\u771f\u7684\u662f\u4e0d\u5bb9\u6613\u554a...<\/p>\n

\u53c3\u8003\u6587\u737b\uff1a
\n<\/strong>Myat T. Lin,Alessandro Occhialini,P. John Andralojc,Martin A. J. Parry & Maureen R. Hanson. 2014.\u00a0A faster Rubisco with potential to increase photosynthesis in crops<\/a>. Nature. doi:10.1038\/nature13776
\nSpencer M. Whitney, Robert L. Houtz and Hernan Alonso. 2011.\u00a0Advancing Our Understanding and Capacity to Engineer Nature\u2019s CO2-Sequestering Enzyme, Rubisco<\/a>. Plant Physiol. 155:27-35<\/p>\n

--
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